以宽川铺生物群中获取的磷酸盐化立体保存的胚胎化石为研究材料，对胚胎化石表面超微结构和内部解剖结构进行观察并对比分析。结果显示，大部分胚胎化石表面被一单层光滑卵膜结构所包裹，卵膜之下胚胎形态多样；刺状结构位于卵膜之下，为原生质胚胎分泌而成的保护层，出现在不同大小和不同类型的胚胎中。结果表明：1)单一依靠胚胎大小或者表面结构不能作为胚胎的分类依据；2)与现生海葵等动物胚胎表面微绒毛相似，Olivooides表面刺状结构可能具有保护胚胎和生态扩散的功能；3)带刺 Olivooides很可能发育成为动物演化早期最简单的带刺幼虫形式，是动物间接发育的证据。

Embryos are preserved in 3D as phosphatized permineralizations with superficial structures,and interior anatomy revealed in partly decayed embryos.Different species share a similar naked egg membrane and stellate cuticle which indicate that:1)It is improper to judge affinity by size or superficial structure of embryos; 2) It is postulated that the stellate cuticle would provide physical protection and dispersal to suitable habitat.Such properties were also observed in extant sea anemone embryos;3) The stellate Olivooidesmay grow into a primitive larva,which could be the evidence of indirect development.

国家自然科学基金(Nos．：41472015，41202008，41772010，41621003，41720104002)；科技部青年973(No．2013CB837100)，教育部111计划，陕西省自然科学基金(No．2016JQ4026))；中央高校基金(Nos．：310827171013，41202008，2013G3274021，2014G3272013)资助