[关键词]
[摘要]
文中阐述了Kunmingaspis属模式种在鉴定中所出现的问题,即将难以观察辨认的性状连接关系简单地当作确切的分类依据,造成非规范化的系统描述。指出张文堂(1964)区分Kunmingaspis和Yuehsienszella两属特征显著差别之一的“眼脊内侧由头鞍的前侧角向外伸出”的论据不可靠,应改正为头鞍前叶或头鞍前外缘是否壮大或弯曲状态。Jell和Hughes(1997)延用并根据两属的模糊概念,在没有全面考察King(1941)标本头尾性状组合的情况下,将Ptychoparia sakesarensis归入Yuehsienszella szechuanensis欠妥。文中通过描述采自云南宜良县上伍营龙肇寺东山坡陡坡寺组下部的新材料,分析P. sakesarensis King,1941和Yuehsienszella szechuanensis (Sun in Lu,1939)的形态差别,证明Kunmingaspis yunnanensis Chang,1964为P. sakesarensis晚出异名。故Kunmingaspis属的模式种应更名为Kunmingaspis sakesarensis。文中还简单地论述了早期褶颊虫类分类的相关问题。
[Key word]
[Abstract]
Based on the new material collected from the Douposi Formation at Shangwuying Village, Yiliang County Yunnan Province, the type species of the genus Kunmingaspis Chang, 1964 is revised. We deny the opinion to identify differentiation between two genera (Kunmingaspis and Yuehsienhszella), Zhang (1964) and Jell and Hughes (1997) considered that “the eye ridge meeting the axial furrow at the level of the rear of the anterior glabellar lobe is distinctive of the genus”. Because an indistinct connection between frontal lobe of glabella and ocular ridge is difficult to be identified on some specimens. The real differentiations selected to be valid are the outline and feature of the frontal lobe of the glabella. Similarly, such as tip of organs about characters (also called character of degradation, or obscure character) can not be excessively weighted in stage of early classification or identification when the phylogeny is unclear,for instance, the S4, the posterior end of several axial ring (may be including the terminal axial piece) of pygidium etc. and invariability and stability of characters should be listed in the diagnosis to distinguish it from other genera. Kunmingaspis differs from Yuehsienszella mainly in having wider less tapering forward glabella with broadly rounded anterior margin, narrower fixigenae between palpebral lobes, relatively longer palpebral lobe of about 1/3 glabellar length (sag.), by contrast in Yuehsienhszella the palpebral lobe is about 1/4 glabellar length (sag.). Besides Yuehsienhszella has a shallower anterior border furrow, wider fixigenae between palpebral lobes (tr.), shallower less curving forward posterior border furrow medially and broader less tapering backward pygidial axis. In the present paper Kunmingaspis yunnanensis Chang, 1964, K. qianbeiensis Zhou in Lu et al., 1974, K. songlinensis Lee in Yin et Lee, 1978, K. longa Zhou in Zhang et al., 1980, K. brevica Zhou in Zhang et al., 1980 are considered as synonymous with K. sakesarensisa (King, 1941). Biostratigraphical, it can be correlated closely between the Yangtze Platform and Himalaya Area (Pakistan or Indian subcontinent).
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[基金项目]
国家自然科学基金(41362002)和国家自然科学青年基金(41402011)联合资助